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In outer space, an unmanned, intelligent life-searching NASA space probe, Infinity, is dragged into an intergalactic wormhole and crash-lands on the other side of the galaxy. It lands on the Earth-like alien planet named Malgor, populated by colorful alien beings. Zartog (Jeff Daniels), an evil-minded inhabitant, accidentally discovers how to take manual control of the onboard machinery and uses it to enslave the population. Faced with the possible extinction of Infinity and their budget, the scientists hire multiple chimpanzees as astronauts to regain contact with the probe and retrieve it: technical genius Comet (Zack Shada), lieutenant Luna (Cheryl Hines) and commander Titan (Patrick Warburton). For media attention, the Senator (Stanley Tucci) adds to the team Ham III (Andy Samberg), grandson of Ham, the first chimpanzee in space, who works as a cannonball at a circus. Ham III is uninterested in the mission, but he (along with the other space chimps (except Comet)) is launched into space despite his best efforts to escape the scientist's training facility.